Antigen presentation and peripheral tolerance T cells only recognize - - PowerPoint PPT Presentation

antigen presentation and peripheral tolerance t cells
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Antigen presentation and peripheral tolerance T cells only recognize - - PowerPoint PPT Presentation

Antigen presentation and peripheral tolerance T cells only recognize antigens presented in the MHC context characteristics Caractersticas Molcula que se liga ao antgeno Molecules bound to antigens Imunoglobulina (Ig) Receptor de clulas


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Antigen presentation and peripheral tolerance

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T cells only recognize antigens presented in the MHC context

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Características Molécula que se liga ao antígeno Imunoglobulina (Ig) Receptor de células T(TCR) Moléculas de MHC* Sítio de ligação ao antígeno 3 regiões CDR nas porções variáveis das cadeias leve e pesada 3 regiões CDR nas porções variáveis α e β Fenda para ligação de peptídeo formada pelas cadeias α1 e β1 em classe II e α1 α2 em classe I Tipo de antígeno Macromolecules, organic Molecules, proteins, lipids, Sugars… MHC/peptide complex Peptídeos Antigen nature Linear or conformacional Linear; 2 to 3 aa residues Linear; 8 to 12 aa depending on the MHC molecule Binding affinity Kd 10-7 - 10-11 M; Affinity maturation Kd 10-5 - 10-7 M Kd 10-6 - 10-9 M; durable interaction

characteristics Molecules bound to antigens Immunoglobulin TCR MHC Binding site 3 CDRs in the light and heavy variable chains 3 CDRs in the alpha and beta variable chains Peptide grooves in the α1/β1 chains in II; In α1/α2 chain in I antigen peptides

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T cell receptor complex

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Characteristics II

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Antigen presentation pathways

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Class II pathway

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HLA-DO ensures peptide presentation diversity

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Class I pathway

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Proteoasome and immunoproteasome

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Immunoproteasome cleaves proteins preferentially after basic and hydrophobic residues generating peptides with higher affinity to MHC-I

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CROSS-PRESENTATION how can a DC activate a CD8 T cell response if it is not infected by a virus?

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PROFESSIONAL ANTIGEN PRESENTING CELLS

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Function – antigen presentation

Antigen -

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Location, location, location……

http://www.youtube.com/watch?v=Kh- XdNnTZUo&feature=player_embedded

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  • 1st and 2nd signals for lymphocyte activation, origin of 2nd signals
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  • Activation of T cells

Immunological synapse

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Co-stimulatory molecules family

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co-stimulation B7.1/B7.2 or CD80/CD86 Stabilization of NFAT Activation of

  • ther txn factors

Co-stimulation

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Diehn et al, PNAS, 18, 11796 (2002)

Stabilization of NFAT signaling

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Activation of other transcription factors

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IL-2 and its receptor

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PERIPHERAL TOLERANCE

Tolerance to self or non harmful antigens triggered in the peripheral tissues. 1 - anergy 2- apoptosis (co-stimulus induce the anti-apoptotic protein Bcl-xL) 3 – Regulatory phenotype induction – dependent on the microenvironment

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REGULATORY T CELLS

TGFb, IL-10 kinurenina naturais induzidas

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FUNCTION – MECHANISMS OF ACTION

EXPRESSÃO DE CITOCINAS INIBIDORAS INDUÇÃO DE MORTE CELULAR ALTERAÇÕES METABÓLICAS INDUÇÃO DE TOLERÂNCIA EM DC INIBIÇÃO DE MATURAÇÃO E FUNÇÃO MORTE POR FALTA DE IL-2 mecanismo extrínsico de CTLA4

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Tregs convert ATP in adenosine – a suppressor molecule

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Tregs express high levels of CD25 and quench IL-2 from the environment

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Tregs limit the proliferation rate of effector T cells

https://doi.org/10.3389/fimmu.2018.02461

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Oral tolerance

Tolerance to antigens acquired through the oral and digestive tract/nasal TGFb expression TGFb e IL-10 rich mucosa – Treg inducion

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Indução de T reguladoras na mucosa intestinal

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Células dendríticas produtoras de ácido retinóico também participam do processo de tolerância

  • RAR (receptor de ácido retinóico) liga-se a elementos resposivos no promotor de genes como CCR9 garantindo a

migração de células reguladoras e linfócitos B produtores de IgA para a mucosa;

  • RA induz fenótipo regulador em linfócitos T, cooperando com TGFb;