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Low rate of lineage High rates of diversification lineage diversification Ancestral trait Evolutionary dead Key innovation innovation ends (e.g. hypothesis for specialization diversity hypothesis) Low rate of trait Niche conservatism

  1. Low rate of lineage High rates of diversification lineage diversification Ancestral trait Evolutionary dead Key innovation innovation ends (e.g. hypothesis for specialization diversity hypothesis) Low rate of trait Niche conservatism diversification no special name? Non-adaptive radiation High rate of trait Adaptive Adaptive radiation diversification divergence? Local adaptation?

  2. Harmon et al. 2003 Science

  3. early diversification -> greater phylogenetic signal Harmon 03-3 Harmon et al. 2003 Science

  4. Measuring niche conservatism - phylogenetic signal Blomberg’s K: measures degree of similarity among close relatives, relative to expectations based on Brownian motion convergence brownian conserved K<<1 K~1 K>>1 K: Blomberg et al. (2003) Evolution ; examples: Ackerly, PNAS in review

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